# Selective culinary uses of plant foods by Northern and Eastern European hunter-gatherer-fishers

**Authors:** Lara González Carretero, Alexandre Lucquin, Harry K. Robson, T. Rowan McLaughlin, Ekaterina Dolbunova, Jasmine Lundy, Giulia Moretti, Blandine Courel, Manon Bondetti, Marian Berihuete-Azorín, Daniel Groß, Jacek Kabaciński, Piotr Kittel, Elena Kostyleva, Olga Lozovskaya, Andrey Mazurkevich, Bente Philippsen, Andrey Skorobogatov, Roman V. Smolyaninov, John Meadows, Carl Heron, Oliver E. Craig

PMC · DOI: 10.1371/journal.pone.0342740 · PLOS One · 2026-03-04

## TL;DR

Northern and Eastern European hunter-gatherers selectively used plant foods in pottery, combining them with animal ingredients, as revealed by a multi-method analysis of foodcrusts.

## Contribution

A combined analytical approach reveals selective plant food use in hunter-gatherer diets, beyond what lipid analysis alone can detect.

## Key findings

- Foodcrusts contained plant tissues like wild grasses, legumes, fruits, and herbaceous plant parts.
- Hunter-gatherers selectively used specific plant species and parts, often in combination with animal ingredients.
- Lipid residue analysis alone under-represents the diversity of plant processing in pottery.

## Abstract

Carbonised food deposits preserved in pottery vessels, often termed ‘foodcrusts,’ are frequently encountered on hunter-gatherer-fisher (HGF) pottery throughout Northern and Eastern Europe. While lipid residue analysis is frequently employed to determine their composition, this technique favours the identification of animal products. In this study, we present a combined analytical approach, including high resolution microscopic analysis (Digital Microscopy and Scanning Electron Microscopy) together with molecular and isotopic analysis of lipids (GC-MS and GC-C-IRMS) and bulk isotope analysis (EA-IRMS) to further understand the composition of foodcrusts. Eighty-five pottery vessels with foodcrusts were analysed from 13 archaeological sites dating from the 6th to the 3rd millennium BC, of which 58 have allowed for identification of plant tissues, such as wild grasses and legumes, fruits, and the roots, tubers, leaves and stems of herbaceous plants. The results demonstrate that the choice of plant foods was remarkably selective, with hunter-gatherers favouring certain plant species and even their parts over others and combining these with specific animal ingredients. The results also reveal that our knowledge of plant processing in pottery is likely to be grossly under-represented by relying on lipid residue analysis alone.

## Full-text entities

- **Diseases:** flowering (MESH:C000719190), Beta vulgaris (MESH:D016112), toxic (MESH:D064420)
- **Chemicals:** beta-Amyrin (MESH:C036380), alpha-Amyrin (MESH:C000654244), Lipid (MESH:D008055), sterols (MESH:D013261), C16:0 (-), triterpenoids (MESH:D014315), starch (MESH:D013213), fatty acids (MESH:D005227), carbohydrates (MESH:D002241), terpenoids (MESH:D013729), fish oils (MESH:D005395), methanol (MESH:D000432), N (MESH:D009584), C (MESH:D002244), phytosterols (MESH:D010840)
- **Species:** Oxybasis glauca (oakleaf goosefoot, species) [taxon 244509], Bromus (genus) [taxon 4501], Carya cordiformis (bitternut, species) [taxon 139928], Blitum bonus-henricus (Good King Henry, species) [taxon 122298], Viburnum opulus (crampbark, species) [taxon 85293], Cyprinus carpio (carp, species) [taxon 7962], Gadus morhua (Atlantic cod, species) [taxon 8049], Acer pensylvanicum (goosefoot, species) [taxon 66214], Avena sativa (cultivated oat, species) [taxon 4498], Homo sapiens (human, species) [taxon 9606], Atriplex prostrata (halberd-leaf orache, species) [taxon 190330], Allium ursinum (ramson, species) [taxon 4684], Chenopodium album (common lambsquarters, species) [taxon 3559], Avena strigosa (black oat, species) [taxon 38783], Beta vulgaris subsp. maritima (sea beet, subspecies) [taxon 350892], P. major [taxon 165745], Allium sativum (garlic, species) [taxon 4682], Conopodium majus (species) [taxon 173702], Beta vulgaris (beet, species) [taxon 161934], Plantago lanceolata (narrow-leaved plantain, species) [taxon 39414], Bolboschoenus maritimus (species) [taxon 76417], Sarcobatus vermiculatus (greasewood, species) [taxon 46107]
- **Mutations:** 4300-4000 cal BC

## Full text

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## Figures

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## References

65 references — full list in the complete paper: https://tomesphere.com/paper/PMC12959656/full.md

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Source: https://tomesphere.com/paper/PMC12959656