# Earliest millet cultivation reflects steppe connections, dietary flexibility, and resilience in Bronze Age northern Greece

**Authors:** Kyriaki Karanikola, Soultana-Maria Valamoti, Giedrė Motuzaite Matuzeviciute, Stefanos Gimatzidis, Stefanos Gimatzidis, Stefanos Gimatzidis, Stefanos Gimatzidis

PMC · DOI: 10.1371/journal.pone.0344099 · PLOS One · 2026-03-03

## TL;DR

This study shows that early millet cultivation in Bronze Age northern Greece was influenced by connections to the steppe and reflects regional dietary and cultural differences.

## Contribution

The study provides new evidence on the introduction and significance of millet in northern Greece through archaeobotanical and radiocarbon data.

## Key findings

- Millet was cultivated in northern Greece during the Bronze Age, unlike in southern Greece.
- Millet cultivation may have been linked to exchange networks connecting the region to the Pontic steppe.
- Millet use reflects cultural and dietary distinctions between northern and southern Greece.

## Abstract

This paper explores early broomcorn millet (hereafter millet) cultivation in Greece during the Bronze Age. The primary archaeobotanical data for this study derive from the site of Skala Sotiros on the island of Thasos in northern Greece. The site provides unique insights into localized Bronze Age agricultural practices, revealing both divergence from southern Greece agricultural systems and potential influences from exchange networks that linked northern Greece to the southern Balkans and the Pontic steppe–Black Sea region. Systematic sampling of the Bronze Age layers at Skala Sotiros has yielded a diverse assemblage with a notable abundance of millet (Panicum miliaceum), a crop almost absent from contemporary southern Greece. Recent radiocarbon dates on millet grains from Skala Sotiros contribute new evidence toward understanding the routes through which millet could have been introduced into the region during the Bronze Age. This study explores the interplay of environmental and cultural factors in the dispersal of millet in Greece, considering environmental stress, cultural dynamics, population movements, and interaction networks. The extensive review of archaeobotanical data across Greece demonstrates how the cultivation of millet may have served as a culinary identity signifier, providing further evidence of differences between northern and southern Greece.

## Linked entities

- **Species:** Panicum miliaceum (taxon 4540)

## Full-text entities

- **Diseases:** drought (MESH:C536747), fire (MESH:D000092422), Bitter vetch (MESH:D013651)
- **Chemicals:** LBA (-), Oil (MESH:D009821), AMS (MESH:D000576), oxygen (MESH:D010100), gold (MESH:D006046), carbon (MESH:D002244), 14C (MESH:C000615234), Iron (MESH:D007501), water (MESH:D014867), tin (MESH:D014001)
- **Species:** Lathyrus clymenum (species) [taxon 3855], Alopecurus pratensis (foxtail, species) [taxon 15304], Powellomyces sp. EA (species) [taxon 252690], Homo sapiens (human, species) [taxon 9606], Fagus (beech trees, genus) [taxon 21024], Vicia faba (broad bean, species) [taxon 3906], Equus caballus (domestic horse, species) [taxon 9796], Bos taurus (bovine, species) [taxon 9913], Lathyrus ochrus (yellow-flowered pea, species) [taxon 3858], Sorghum bicolor (broomcorn, species) [taxon 4558], Triticum aestivum (bread wheat, species) [taxon 4565], Hordeum sp. (species) [taxon 50472], Triticum spelta (spelt, species) [taxon 58933], Hordeum vulgare (barley, species) [taxon 4513], Typha latifolia (common cattail, species) [taxon 4733], Olea europaea (common olive, species) [taxon 4146], Triticum timopheevii (Sanduri wheat, species) [taxon 4570], Triticum monococcum (einkorn wheat, species) [taxon 4568], Panicum miliaceum (broomcorn millet, species) [taxon 4540], Olea (olives, genus) [taxon 4145], Avena sp. (oat, species) [taxon 50455], Triticum turgidum subsp. dicoccum (cultivated emmer wheat, subspecies) [taxon 49225], Lathyrus sativus (chickling vetch, species) [taxon 3860], Lens culinaris (lentil, species) [taxon 3864], Vicia ervilia (species) [taxon 154498], Vitis vinifera (wine grape, species) [taxon 29760], Buglossoides arvensis (corn gromwell, species) [taxon 181181]

## Full text

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## Figures

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## References

121 references — full list in the complete paper: https://tomesphere.com/paper/PMC12956113/full.md

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Source: https://tomesphere.com/paper/PMC12956113