# Morphological and Phylogenetic Evidence Reveal Nine New Species of Russula (Russulaceae, Russulales) from Shanxi Province, North China

**Authors:** Hao-Yu Fu, Jia-He Li, Hui-Min Ji, Ning Mao, Ting Li, Li Fan

PMC · DOI: 10.3390/jof12010078 · Journal of Fungi · 2026-01-22

## TL;DR

This paper reports nine new species of the mushroom genus Russula found in Shanxi Province, northern China, using both physical traits and genetic analysis.

## Contribution

The discovery of nine new Russula species in northern China using combined morphological and phylogenetic methods.

## Key findings

- Nine new Russula species were identified in Shanxi Province using morphological and genetic analyses.
- The new species are distributed across three subgenera: Brevipes, Heterophyllidia, and Russula.
- The study improves understanding of Russula diversity in China's temperate regions.

## Abstract

Shanxi Province, located in northern China, characterized by a warm-temperate monsoon climate, complex mountainous topography, and vegetation dominated by trees of Fagaceae and Pinaceae, provides diverse habitats for Russula diversity. Recent investigations on macrofungi in this region revealed nine new Russula species based on integrated morphological and multi-locus phylogenetic analyses (ITS, nrLSU, rpb2, tef1), which are described and illustrated in this paper. These new taxa are classified into three subgenera of Russula: one species of subgen. Brevipes, four of subgen. Heterophyllidia, four of subgen. Russula. This work enhances the understanding of Russula resources in China’s temperate zone.

## Linked entities

- **Species:** Russula (taxon 5402), Fagaceae (taxon 3503), Pinaceae (taxon 3318)

## Full-text entities

- **Species:** Russula (genus) [taxon 5402]

## Full text

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## Figures

22 figures with captions in the complete paper: https://tomesphere.com/paper/PMC12843102/full.md

## References

74 references — full list in the complete paper: https://tomesphere.com/paper/PMC12843102/full.md

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Source: https://tomesphere.com/paper/PMC12843102