# Systematics Review and Phylogeny of Cyrtophyllitinae Zeuner, 1935 sensu Gorochov, Jarzembowski & Coram, 2006 (Ensifera, Haglidae), with Description of Two New Species

## TL;DR

This paper revises the classification of a group of insects called Cyrtophyllitinae, describing two new species and a new subfamily from Middle Jurassic China.

## Contribution

The paper introduces a new subfamily and genus, and describes two new species based on Middle Jurassic fossils from China.

## Key findings

- Cyrtophyllitinae is found to be paraphyletic, leading to the creation of a new subfamily Archaboilinae.
- Two new species, Archaboilus ornatus and Vitimoilus gigantus, were identified from Middle Jurassic China.
- A new genus, Pararchaboilus, was established with a designated type species.

## Abstract

The systematics of Cyrtophyllitinae Zeuner, 1935 sensu Gorochov, Jarzembowski & Coram, 2006 was revised. A new subfamily Archaboilinae subfam. nov. and a new genus, Pararchaboilus gen. nov. were erected. Two new species, Archaboilus ornatus sp. nov. and Vitimoilus gigantus sp. nov., are described from the Middle Jurassic China.

A phylogeny of Cyrtophyllitinae Zeuner, 1935 sensu Gorochov, Jarzembowski & Coram, 2006, based on wing morphology, is presented including all genera. Cyrtophillitinae is found to be paraphyletic. Except for Cyrtophyllites rogeri Oppenheim, 1888, all other species were moved from the subfamily Cyrtophyllitinae (Hagloidea, Haglidae). Consequently, a new subfamily Archaboilinae subfam. nov. was erected and accommodates most of the previous cyrtophillitine taxa, except Cyrtophyllites rogeri. The type genus Archaboilus Martynov, 1937 of the new subfamily was designated; a new genus, Pararchaboilus gen. nov., was erected with the designation of type species Pararchaboilus cretaceus comb. nov. From the Middle Jurassic deposits of China, two new species, Archaboilus ornatus sp. nov. and Vitimoilus gigantus sp. nov., are described.