Symbiotic behaviour in the Public Goods game with altruistic punishment
Lucas S. Flores, Heitor C. M. Fernandes, Marco A. Amaral and, Mendeli H. Vainstein

TL;DR
This study introduces altruistic punishers in a spatial public goods game, revealing that their symbiotic relationship with cooperators can sustain cooperation and punish defectors, especially in structured populations with certain costs.
Contribution
It demonstrates a novel symbiotic mechanism where altruistic punishers and cooperators coexist to promote cooperation in spatial public goods games.
Findings
Punishers thrive by sharing costs at low punishment levels.
Cooperators are driven to extinction without punishers.
Symbiotic spatial structures emerge at higher punishment costs.
Abstract
Finding ways to overcome the temptation to exploit one another is still a challenge in behavioural sciences. In the framework of evolutionary game theory, punishing strategies are frequently used to promote cooperation in competitive environments. Here, we introduce altruistic punishers in the spatial public goods game. This strategy acts as a cooperator in the absence of defectors, otherwise it will punish all defectors in their vicinity while bearing a cost to do so. We observe three distinct behaviours in our model: i) in the absence of punishers, cooperators (who don't punish defectors) are driven to extinction by defectors for most parameter values; ii) clusters of punishers thrive by sharing the punishment costs when these are low iii) for higher punishment costs, punishers, when alone, are subject to exploitation but in the presence of cooperators can form a symbiotic spatial…
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