Propagation dynamics of Fisher-KPP equation with time delay and free boundaries
Ningkui Sun, Jian Fang

TL;DR
This paper studies how time delay and free boundaries influence the propagation of solutions in Fisher-KPP equations, revealing a dichotomy between spreading and vanishing, and showing delays slow down spreading speed.
Contribution
It establishes a well-posedness condition for delayed reaction-diffusion equations with free boundaries and characterizes the spreading speed and profile affected by delays.
Findings
A vanishing-spreading dichotomy is proven.
Spreading speed is slowed by time delay.
Spreading profile is determined by a nonlocal semi-wave problem.
Abstract
Incorporating free boundary into time-delayed reaction-diffusion equations yields a compatible condition that guarantees the well-posedness of the initial value problem. With the KPP type nonlinearity we then establish a vanishing-spreading dichotomy result. Further, when the spreading happens, we show that the spreading speed and spreading profile are nonlinearly determined by a delay-induced nonlocal semi-wave problem. It turns out that time delay slows down the spreading speed.
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Taxonomy
TopicsMathematical and Theoretical Epidemiology and Ecology Models · Differential Equations and Numerical Methods · Numerical methods for differential equations
Propagation dynamics of Fisher-KPP equation with time delay and free boundaries
Ningkui Sun
School of Mathematics and Statistics
Shandong Normal University
Jinan 250014, China
and
Jian Fang
Institute for Advanced Studies in Mathematics and Department of Mathematics
Harbin Institute of Technology
Harbin 150001, China
Abstract.
Incorporating free boundary into time-delayed reaction-diffusion equations yields a compatible condition that guarantees the well-posedness of the initial value problem. With the KPP type nonlinearity we then establish a vanishing-spreading dichotomy result. Further, when the spreading happens, we show that the spreading speed and spreading profile are nonlinearly determined by a delay-induced nonlocal semi-wave problem. It turns out that time delay slows down the spreading speed.
Key words and phrases:
reaction-diffusion equation, free boundary, time delay, spreading phenomena
2010 Mathematics Subject Classification:
35K57, 35R35, 35B40, 92D25
The research leading to these results was supported by the NSF of China (No.11771108), the NSF of Heilongjiang province of China (No. LC2017002) and the NSF of Shandong Province of China (No. ZR201702140024).
1. Introduction
In the pioneer work of Fisher [17], and Kolmogorov, Petrovski and Piskunov [22], it was shown that
[TABLE]
with
[TABLE]
admits traveling waves solutions of the form satisfying and if and only if . In 1970s’, Aronson and Weinberger [2] proved that the minimal wave speed is also the asymptotic speed of spread (spreading speed for short) in the sense that
[TABLE]
for any small provided that the initial function is compactly supported. These works have stimulated volumes of studies for the propagation dynamics of various types of evolution systems. Among others, of particular interest to the Fisher-KPP equation (1.1)-(1.2) with time delay or free boundary are two typical ones.
Schaaf [32] studied the following delayed reaction-diffusion equation
[TABLE]
where is the time delay. With the Fisher-KPP condition on and the quasi-monotone condition , it was shown that the minimal wave speed exists and it is determined by the system of two transcendental equations
[TABLE]
where
[TABLE]
The delay-induced spatial non-locality was brought to attention by So, Wu and Zou [28], where they derived the following time-delayed reaction-diffusion model equation with nonlocal response for the study of age-structured population
[TABLE]
where represents the density of mature population, is the maturation age, is the death rate, is the birth rate function, is the survival rate from newborn to being mature, and is the redistribution kernel during the maturation period. As such, introducing time delay into diffusive equation usually gives rises to spatial non-locality due to the interaction of time lag (for maturation) and diffusion of immature population. In the extreme case where the immature population does not diffuse, the kernel becomes the Dirac measure, and hence (1.7) reduces to (1.4). We refer to the survey article [21] for the delay-induced nonlocal reaction-diffusion problems. In [28], the authors obtained the minimal wave speed that is determined by a similar system to (1.5) provided that is nondecreasing and is of Fisher-KPP type. Wang, Li and Ruan [37] proved that is decreasing in . Liang and Zhao [23] showed that is also the spreading speed for the solutions satisfying the following initial condition
[TABLE]
Similar to the classical Fisher-KPP equation, the spreading speed for delayed reaction-diffusion equation is still linearly determined for both local and nonlocal problems thanks to the Fisher-KPP type condition.
We refer to [26] for more properties that are induced by time delay in reaction-diffusion equations, including the well-posedness of initial value problems as well as the role of the quasi-monotone condition on the comparison principle, and [14, 15] for the delay-induced weak compactness of time- solution maps when as well as its role in the study of wave propagation.
Recently, Du and Lin [10] proposed a Stefan type free boundary to the Fisher-KPP equation
[TABLE]
where the free boundaries and represent the spreading fronts, which are determined jointly by the gradient at the fronts and the coefficient in the Stefan condition. For more background of proposing such free boundary conditions, we refer to [10, 7]. It was proved in [10] that the unique global solution has a spreading-vanishing dichotomy property as : either and (spreading case), or , with , and (vanishing case). Moreover, it was also proved that when spreading happens, there is a constant such that and behave like a straight line for large time, where is called the asymptotic speed of spread (spreading speed for short). Different from the classical Fisher-KPP speed, is the unique value of such that the following nonlinear semi-line problem is solvable:
[TABLE]
where is the right derivative of at [math]. In particular, as increases to infinity, increases to the classical Fisher-KPP speed . Later on, Du and Lou [11] obtained a rather complete characterization on the asymptotic behavior of solutions for (1.9) with some general nonlinear terms. For further related work on free boundary problems, we refer to [8, 9, 12] and the references therein.
In this paper, we aim to explore how to incorporate time delay and free boundary into the Fisher-KPP equation (1.1)-(1.2) so that the problem is well-posed, and then study their joint influence on the propagation dynamics.
Keeping a smooth flow for the organizations of the paper, we write down here the problem of interest while leaving in the next section the derivation details, including the emergence of the compatible condition (1.12) for the well-posedness of the initial value problem.
[TABLE]
where and are two positive constants, the nonlinear function satisfies
[TABLE]
and the initial data satisfies
[TABLE]
as well as the compatible condition
[TABLE]
Assumption (H) ensures the Fisher-KPP structure as well as the comparison principle. Due to the nature of delay differential equations, the initial value, including the initial domain, has to be imposed over the history period , as in (1.11). The interaction of time delay and free boundary gives rise to the compatible condition (1.12) that is essential for the well-posedness of the problem. If , then the compatible condition (1.12) becomes trivial and problem (P) reduces to (1.9).
Theorem 1.1**.**
(Well-posedness)* For an initial data satisfying (1.11) and (1.12), there exists a unique triple solving () with and .*
With the compatible condition (1.12) we can cast the problem into a fixed boundary problem and then apply the Schauder fixed point theorem to establish the local existence of solutions. The extension to all positive time is based on some a priori estimates111We sincerely thank Professor Avner Friedman for his valuable comments and suggestions on the proof of the well-posedness..
From the maximum principle and (H), it follows that when the solution as , and , and hence, for all . Therefore, we can denote
[TABLE]
Theorem 1.2**.**
(Spreading-vanishing dichotomy)* Let be the solution of (). Then the following alternative holds:*
Either
(i) Spreading:* and*
[TABLE]
or
(ii) Vanishing:* is a finite interval with length no bigger than and*
[TABLE]
When spreading happens, we characterize the spreading speed and profile of the solutions. The nonlinear and nonlocal semi-wave problem
[TABLE]
will play an important role. If then (1.13) reduces to the local form (1.10).
Theorem 1.3**.**
Problem (1.13) admits a unique solution and is decreasing in delay .
Due to the presence of time delay, the proof of Theorem 1.3 highly relies on the distribution of complex solutions of the following transcendental equation
[TABLE]
We refer to Lemma 3.2 and Proposition 3.3, which are independently of interest.
With the semi-wave established above, we can construct various super- and subsolutions to estimate the spreading fronts and the spreading profile as .
Theorem 1.4**.**
(Spreading profile)* Let be a solution satisfying Theorem 1.2(i). Then there exist two constants and such that*
[TABLE]
[TABLE]
[TABLE]
[TABLE]
where is the unique solution of (1.13).
The rest of this paper is organized as follows. In Section 2 we derive the compatible condition (1.12), with which we formulate problem (P) and then establish the well-posedness as well as the comparison principle. Section 3 is devoted to the study of the semi-wave problem (1.13). In section 4, we establish the spreading-vanishing dichotomy result. Finally in Section 5, we characterize the spreading speed and profile of spreading solutions of ().
2. The compatible condition, well-posedness and comparison principle
2.1. The compatible condition
To formulate problem (), we start from the age-structured population growth law
[TABLE]
where denotes the density of species of age at time and location , and denote the diffusion rate and death rate of species of age , respectively.
Next we consider the scenario that the species has the following biological characteristics.
- (A1)
The species can be classified into two stages by age: mature and immature. An individual at time belongs to the mature class if and only if its age exceeds the maturation time . Within each stage, all individuals share the same behavior.
- (A2)
Immature population does not move in space.
The total mature population at time and location can be represented by the integral
[TABLE]
We assume that the mature population lives in the habitat , vanishes in the boundary
[TABLE]
and extends the habitat by obeying the Stefan type moving boundary conditions:
[TABLE]
where is a given positive constant. Note that the immature population does not contribute to the extension of habitat due to their immobility, as assumed in (A2).
According to (A1) we may assume that
[TABLE]
where and are two positive constants. Differentiating the both sides of the equation (2.2) in time yields
[TABLE]
Since no individual lives forever, it is nature to assume that
[TABLE]
To obtain a closed form of the model, one then needs to express by in a certain way. Indeed, denotes the newly matured population at time , and it is the evolution result of newborns at . In other words, there is an evolution relation between the quantities and . Such a relation is obeyed by the growth law (2.1) for , and hence it is the time- solution map of the following equation
[TABLE]
Therefore, . Further, the newborns is given by the birth , where is the birth rate function with . Consequently,
[TABLE]
Combining (2.3)-(2.6) and (2.8), we are led to the following system:
[TABLE]
For , outside the habitat the mature population does not exist, that is,
[TABLE]
Clearly, since the habitat is expanding for , we have
[TABLE]
Hence, the first two equations in (2.9) can be written as the following single one
[TABLE]
provided that (2.11) holds for . As such, in view of (2.11) we need an additional condition
[TABLE]
Note that for . And as the coefficient we have uniformly for . Therefore, regardless of the influence of , (2.13) is strengthened to be
[TABLE]
which is the aforementioned compatible condition (1.12).
Setting in (2.9), we obtain problem ().
2.2. Well-posedness
We employ the Schauder fixed point theorem to establish the local existence of solutions to (), and prove the uniqueness, then extend the solutions to all time by an estimate on the free boundary.
Theorem 2.1**.**
Suppose (H) holds. For any , there is a such that problem () admits a solution
[TABLE]
Proof.
We divide the proof into three steps.
. We use a change of variable argument to transform problem () into a fixed boundary problem with a more complicated equation which is used in [3, 10]. Denote and for convenience, and set . Let and be two nonnegative functions in such that
[TABLE]
[TABLE]
Define through the identity
[TABLE]
and set
[TABLE]
Then the free boundary problem () becomes
[TABLE]
and
[TABLE]
with and ,
[TABLE]
Denote , and . For 0<T\leqslant\min\big{\{}\frac{h_{0}}{16(1+h_{1}+h_{2})},\ \tau\big{\}}, we define ,
[TABLE]
Clearly, is a bounded and closed convex set of .
Noting that the restriction on , it is easy to see that the transformation is well defined. By a similar argument as in [36], applying standard theory and the Sobolev embedding theorem, we can deduce that for any given , problem (2.14) admits a unique , which satisfies
[TABLE]
where and is a constant dependent on , , , and .
Defining and by and , respectively, then we have
[TABLE]
and thus , which satisfies
[TABLE]
Similarly , which satisfies
[TABLE]
. For any given triple , we define an operator by
[TABLE]
Clearly, is continuous in , and is a fixed point of if and only if solves (2.14) and (2.15). We will show that if is small enough, then has a fixed point by using the Schauder fixed point theorem.
Firstly, it follows from (2.17) and (2.18) that
[TABLE]
Thus if we choose T\leqslant\min\big{\{}\frac{h_{0}}{16(1+h_{1}+h_{2})},\ \tau,\ C^{-\frac{2}{\alpha}}_{2}\big{\}}, then maps into itself. Consequently, has at least one fixed point by using the Schauder fixed point theorem, which implies that (2.14) and (2.15) have at least one solution defined in . Moreover, by the Schauder estimates, we have additional regularity for as a solution of (2.14) and (2.15), namely,
[TABLE]
and for any given , there holds
[TABLE]
where is a constant dependent on , , , and . Thus we deduce a local classical solution of () by , and satisfies
[TABLE]
. We will prove the uniqueness of solutions of (). Let , , be two solutions of () and set
[TABLE]
Then it follows from (2.16), (2.17) and (2.18) that
[TABLE]
Set
[TABLE]
then we find that satisfies that
[TABLE]
where
[TABLE]
and and are the coefficients of problem (2.14) with instead of .
Recalling that , then for all , thus
[TABLE]
Thanks to this, we can apply the estimates for parabolic equations to deduce that
[TABLE]
with depending on and . By a similar argument as in [36], we obtain that
[TABLE]
for some positive constant independent of . Thus
[TABLE]
Since , then
[TABLE]
This, together with (2.21), implies that
[TABLE]
where . Similarly, we have
[TABLE]
As a consequence, we deduce that
[TABLE]
Hence for
[TABLE]
we have
[TABLE]
This shows that for , thus in . Consequently, the uniqueness of solution of () is established, which ends the proof of this theorem. ∎
Lemma 2.2**.**
Assume that (H) holds. Then every positive solution of problem () exists and is unique for all .
Proof.
Let be the maximal time interval in which the solution exists. In view of Theorem 2.1, it remains to show that . We proceed by a contradiction argument and assume that . Thanks to the choice of the initial data, the comparison principle implies that for . Construct the auxiliary function
[TABLE]
where
[TABLE]
It follows the proof of [10, Lemma 2.2] to prove that there is a constant independent on such that for . The proof for for is parallel.
Let us now fix . Similar to the proof of Theorem 2.1, by standard estimate, the Sobolev embedding theorem and the Hölder estimates for parabolic equation, we can find depending only on , , , , and such that
[TABLE]
This implies that exists on . Choosing with , and regarding for as the initial function, it then follows from the proof of Theorem 2.1 that there exists depending on , and independent of such that problem () has a unique solution in . This yields that the solution of () can be extended uniquely to . Hence when is large. But this contradicts the assumption, which ends the proof of this lemma. ∎
Proof of Theorem 1.1: Combining Theorem 2.1 and Lemma 2.2, we complete the proof.
2.3. Comparison Principle
In this subsection, we establish the comparison principle, which will be used in the rest of this paper. Let us start with the following result.
Lemma 2.3**.**
Suppose that (H) holds, , , satisfies in with , and
[TABLE]
If for and satisfies
[TABLE]
then the solution of problem () satisfies , in , and
[TABLE]
Proof.
We integrate the ideas of [10, Lemma 5.7] and [26, Corollary 5] to deal with free boundary and time delay.
Firstly, for small , let denote the unique solution of () with and replaced by and for , respectively, with replaced by , and with replaced by some , satisfying
[TABLE]
and for any fixed as , in the norm.
We claim that , and for all and . Obviously, this is true for all small . Now, let us use an indirect argument and suppose that the claim does not hold, then there exists a first such that
[TABLE]
and there is some such that .
Later, let us compare and over the region
[TABLE]
An direct computation shows that for ,
[TABLE]
it then follows from the strong maximum principle that
[TABLE]
Thus either or . Without loss of generality we may assume that , then . This, together with (2.23), implies that , from which we obtain that
[TABLE]
As for and , then , which contradicts (2.24). This proves our claim.
Finally, thanks to the unique solution of () depending continuously on the parameters in (), as , converges to , the unique of solution of (). The desired result then follows by letting in the inequalities and . ∎
By slightly modifying the proof of Lemma 2.3, we obtain a variant of Lemma 2.3.
Lemma 2.4**.**
Suppose that (H) holds, , , satisfies in with , and
[TABLE]
with in , , for and , where is a solution to (). Then
[TABLE]
Remark 2.5**.**
The function , or the triple , in Lemmas 2.3 and 2.4 is often called a supersolution to (). A subsolution can be defined analogously by reversing all the inequalities. There is a symmetric version of Lemma 2.4, where the conditions on the left and right boundaries are interchanged. We also have corresponding comparison results for lower solutions in each case.
3. Semi-waves
This section is devoted to proving the existence and uniqueness of a semi-wave of (1.13), which will be used to construct some suitable sub- and supersolutions to study the asymptotic profiles of spreading solutions of (). Let us consider the following nonlocal elliptic problem
[TABLE]
where is a constant.
If is understood as the time variable, then we may regard problem (3.1) as a time-delayed dynamical system in the phase space . When , the phase space reduces to and it follows from the phase plane analysis that (3.1) admits a unique positive solution , which is increasing in and as . When , the phase space is of infinite dimension and the positivity and boundedness of the unique solution are not clear.
Proposition 3.1**.**
Suppose (H) holds. For any given constant , problem (3.1) has a maximal nonnegative solution . Moreover, either or in . Furthermore, if , then it is the unique positive solution of (3.1), in and as , in addition, for any given constant , one has for , and .
Proof.
We divide the proof into four steps.
. Problem (3.1) always has a maximal nonnegative solution and it satisfies
[TABLE]
Clearly, [math] is a nonnegative solution of (3.1). For any , consider the following problem:
[TABLE]
It is well known problem (3.2) admits a unique solution for . Applying the maximal principle, we can deduce that for . Moreover, it is easy to check that is decreasing in and increasing in and
[TABLE]
where is a nonnegative solution of problem (3.1) and it satisfies for .
In what follows, we want to prove that is the maximal nonnegative solution of (3.1). Let be an arbitrary nonnegative solution of (3.1), then for . If , then . Suppose now , then in . Let us show for .
Firstly, for any fixed we can find large such that for . We claim that the above inequality holds for . On the contrary, define
[TABLE]
then and for . Thanks to the monotonicity of in , then there is such that and for . It is easy to check that . Then the strong maximal principle yields that and for . Thus we can find a constant such that
[TABLE]
and for . So there is such that and for .
Later, we want to prove that for all . Combining the definition of , we only need to prove for all . Since for and , and for ,
[TABLE]
where the monotonicity of in and the fact where for are used. The comparison principle yields that for all . This, together with the definition of and (3.3), yields that for all , which contradicts the definition of . Thus we have proved that for .
Finally, letting , we deduce that
[TABLE]
as we wanted. Thus Step 1 is proved.
. For any , if is a positive solution of (3.1), then , for , and as .
Since for , then the Hopf lemma can be used to deduce , it follows that for all small . Setting
[TABLE]
In the following, we shall show . Suppose by way of contradiction that , then
[TABLE]
Define for , then
[TABLE]
Let us set
[TABLE]
Then for and it satisfies
[TABLE]
The strong maximal principle and the Hopf lemma imply that
[TABLE]
It follows the continuity that for all small ,
[TABLE]
which implies that for . Moreover, since , it then follows that . But these facts contradict the definition of . Thus the monotonicity of positive solutions of (3.1) is established.
Next, we consider the asymptotic behavior of positive solution of (3.1). The monotonicity of implies that there is a constant such that . We claim that . For any sequence with as , define . Then solves the same equation as but over . Since , it follows that there is a subsequence of (still denoted by ) such that locally in as , and is a solution of
[TABLE]
On the other hand, it follows from that , which implies that , as we wanted. Thus this completes the proof of Step 2.
. We show that problem (3.1) has at most one positive solution.
Suppose problem (3.1) has two positive solutions and , then in , and as for . Define
[TABLE]
From Step 2 we have , . Then by L’Hôpital’s rule we obtain , which together with implies that . Next we show . Indeed, assume for the sake of contraction that . Define
[TABLE]
Then for , , and
[TABLE]
where the sub-linearity and monotonicity of for are used. By Hopf’s lemma, we see that , which implies that . Thus, in view of the definition of , we have an such that . By the elliptic strong maximum principle, we infer that for , a contradiction to . Therefore, , and hence, . Changing the role of and and repeating the above arguments, we obtain . The uniqueness is proved.
. Let us consider the monotonicity of positive solutions in .
Assume that is a positive solution of (3.1). Choose and let be the maximal nonnegative solution of (3.1) with . Since is a supersolution of (3.1), and by Step 2 we know that is a subsolution of (3.1) with , in view of the uniqueness of positive solution of this problem, then we see that for . It thus follows from the maximum principle and the Hopf lemma that
[TABLE]
The proof of this proposition is complete now. ∎
Next we give a necessary and sufficient condition for the existence of a positive solution of (3.1). For this purpose, we need the following property on the distribution of complex solutions to a transcendental equation.
Lemma 3.2**.**
Let and . Define
[TABLE]
Then there exists such that the following statements hold:
- (i)
* has a positive solution if and only if ;* 2. (ii)
* has a complex solution in the domain*
[TABLE]
provided that .
Before the proof, we note that if then reduces to a polynomial equation of order . It admits at least one positive solution if and only if and exactly a pair of complex eigenvalues in when .
Proof.
(i) Note that is convex in , decreasing in when , and is negative for some when is sufficiently large. Therefore, such exists.
(ii) We employ a continuation method with being the parameter. From the proof of [31, Theorem 2.1], we can infer that the solutions of is continuous in . We write , where and are continuous in . Separating the real and imaginary parts of yields
[TABLE]
We proceed with four steps.
. If is small enough, then there is a solution in . Indeed, At , (3.7) admits a solution . Note that
[TABLE]
It then follows from the implicit function theorem that for small , admits a complex solution near , and hence, in the open domain .
. For any , admits no solution with or when . It follows from statement (i) that there is no solution with when . If equals , then from the second equation of (3.7) we can infer that . Substituting and into the first equation of (3.7), we obtain , a contradiction.
. If a solution touches pure imaginary axis at some , then . We use the implicit function theorem. By direct computations, we have
[TABLE]
where the equality holds if and only if and . Taking these two relations into (3.7) with , we obtain
[TABLE]
which is not solvable for . Therefore,
[TABLE]
On the other hand,
[TABLE]
Consequently, by the implicit function theorem we have
[TABLE]
from which we compute to have
[TABLE]
. Completion of the proof. In Steps 2 and 3, we have verified that the perturbed solution at Step 1 can not escape continuously as increases from [math] to . Therefore, it always stays in . ∎
Based on the above results, we are ready to give the following necessary and sufficient condition for (3.1) to have a unique positive solution.
Proposition 3.3**.**
Suppose (H) holds. Problem (3.1) has a unique positive solution if and only if , where is given in Lemma 3.2.
Proof.
Firstly, let us show that problem (3.1) admits a unique positive solution when . We employ the super- and subsolution method. The case where is trivial and the proof is omitted. Fix . By Lemma 3.2 we can infer that there exists such that
[TABLE]
has a solution in .
Claim. The function
[TABLE]
is a subsolution provided that is small enough.
Indeed, for , we have
[TABLE]
Choose sufficiently small such that
[TABLE]
with which we obtain
[TABLE]
Clearly, if , then , and hence, . If , then , and hence,
[TABLE]
with . Since (as proved in Lemma 3.2), we obtain when . To summarize, for and for . The claim is proved.
Having such a subsolution, we can infer that (3.1) admits a positive solution. The proof of uniqueness of the solution of (3.1) follows from Proposition 3.1.
Next we show that (3.1) does not admit a positive solution when . We employ a sliding argument. Assume for the sake of contradiction that there is a solution . Since , admits a positive solution . Define . Since and , we may choose such that for and vanishes at some . Note that . It then follows that
[TABLE]
By the elliptic strong maximum principle, we obtain for , a contradiction. The nonexistence is proved. ∎
Based on the above results, we obtain the solvability of (1.13).
Theorem 3.4**.**
For any given , let be given in Lemma 3.2. For each , there exists a unique such that , where is the unique positive solution of (3.1) with replaced by . Moreover, is increasing in with
[TABLE]
Proof.
From Propositions 3.1 and 3.3, it is known that for each , problem (3.1) admits a unique solution for , and for any , in . Define
[TABLE]
Then for all and it decreases continuously in . Let . For each problem (3.1) admits a unique solution . Clearly, converges to some and converges to locally uniformly in , and solves (3.1) with . By the nonexistence established in Proposition 3.3 we obtain . In particular,
[TABLE]
We now consider the continuous function
[TABLE]
By the above discussion we know that is strictly decreasing in . Moreover, and . Thus there exists a unique such that , which means that
[TABLE]
Next, let us view as the unique intersection point of the decreasing curve with the increasing line in the -plane, then it is clear that increases to as increases to . The proof is complete. ∎
Remark 3.5**.**
In [11], the authors considered the case . They obtained that for each , there is a unique such that , where is the unique of (3.1) with and , and . Moreover, is increasing in with
[TABLE]
In the rest of this part, we study the monotonicity of in . For any given , the unique positive solution of (3.1) with may be denoted by . Now we give the proof of Theorem 1.3.
Proof of Theorem 1.3: For and , let be given in Theorem 3.4 and Remark 3.5 for and , respectively. By Propositions 3.1 and 3.3, we see that for and , problem (3.1) admits a unique positive solution . Moreover, is increasing in and decreasing in . Let be defined as in (3.14).
Claim. For , when .
We postpone the proof of the claim and reach the conclusion in a few lines. Note that is the unique positive solution of . In view of , we have . If , then we are done. Otherwise, , which, together with the claim, implies that
[TABLE]
This further implies that , due to the monotonicity of in . Thus, is decreasing in .
Proof of the claim. Since is decreasing in , we see that is well-defined when . By the monotonicity of in , we have . This, together with the monotonicity of in , implies that . Consequently,
[TABLE]
Consider the initial value problem
[TABLE]
By the maximum principle we know that is nondecreasing in and its limit as satisfies (3.1) with . By the uniqueness established in Proposition 3.1, we obtain . Therefore,
[TABLE]
The claim is proved.
4. Long time behavior of the solutions
In this section we study the asymptotic behavior of solutions of (). Firstly, we give some sufficient conditions for vanishing and spreading. Next, based on these results, we prove the spreading-vanishing dichotomy result of (). Let us start this section with the following equivalent conditions for vanishing.
Lemma 4.1**.**
Assume that (H) holds. Let be a solution of (). Then the following three assertions are equivalent:
[TABLE]
Proof.
“(i) (ii)”. Without loss of generality we assume and prove (ii) by contradiction. Assume that , then there exists such that
[TABLE]
Let us consider the following auxiliary problem:
[TABLE]
It is easy to check that is a subsolution of (), then and by our assumption. Using a similar argument as in [9, Lemma 3.3] one can show that
[TABLE]
where is the unique positive solution of the problem
[TABLE]
Thus,
[TABLE]
for some , which contradicts the fact that .
“(ii)(iii)”. It follows from the assumption and [39, Proposition 2.9] that the unique positive solution of the following problem
[TABLE]
with in , satisfies uniformly for as . Then the conclusion (iii) follows easily from the comparison principle.
“(iii)(ii)”: Suppose by way of contraction argument that for some small there exists such that for all . Let , it is well known that the following eigenvalue problem
[TABLE]
has a negative principal eigenvalue, denoted by , whose corresponding positive eigenfunction, denoted by , can be chosen positive and normalized by . Set
[TABLE]
with small such that
[TABLE]
It is easy to compute that for ,
[TABLE]
Moreover one can see that
[TABLE]
provided that is sufficiently small. Then we can apply the comparison principle to deduce
[TABLE]
contradicting (iii).
“(ii)(i)”. When (ii) holds, (i) is obvious. This proves the lemma. ∎
Next, we give a sufficient condition for vanishing, which indicates that if the initial domain and initial function are both small, then the species dies out eventually in the environment.
Lemma 4.2**.**
Assume that (H) holds. Let be a solution of (). Then vanishing happens provided that and is sufficient small.
Proof.
Set
[TABLE]
then , so there exists a small such that
[TABLE]
For such , we can find a small positive constant such that
[TABLE]
Define
[TABLE]
and extend by [math] for , .
A direct calculation shows that for ,
[TABLE]
where we have used , for and for .
When and , it is easy to check that
[TABLE]
where the fact that \cos\Big{(}\frac{\pi x}{2k(t-\tau)}\Big{)}\leqslant\cos\Big{(}\frac{\pi x}{2k(t)}\Big{)} for and the monotonicity of in are used. If and , we have that
[TABLE]
Thus we have
[TABLE]
On the other hand,
[TABLE]
As a consequence, will be a supersolution of () if in . Indeed, choose , which depends only on and . Then when we have in , since . It follows from the comparison principle that
[TABLE]
This, together with the previous lemma, implies that vanishing happens. ∎
Remark 4.3**.**
When , the proof of Lemma 4.2 reduces to that of [11, Theorem 3.2(i)].
We now present a sufficient condition for spreading, which reads as follows.
Lemma 4.4**.**
Assume that (H) holds. If , then spreading happens for every positive solution of ().
Proof.
Since for , we have for any . So the conclusion follows from Lemma 4.1. In what follows we prove
[TABLE]
First, it is well known that for any , the following problem
[TABLE]
admits a unique positive solution , which is increasing in and satisfies
[TABLE]
Moreover we can find an increasing sequence of positive numbers with as such that for all . Since converges to locally uniformly in , we can choose such that and for . It then follows from [39] the following problem
[TABLE]
has a unique positive solution , which satisfies that
[TABLE]
Applying the comparison principle we have for all , . This, together with (4.5), yields that
[TABLE]
Later, since the initial data satisfies for , it thus follows from the comparison principle that
[TABLE]
Combining with (4.6), one can easily obtain (4.4), which ends the proof of this lemma. ∎
Now we are ready to give the proof of Theorem 1.2.
Proof of Theorem 1.2. It is easy to see that there are two possibilities: (i) ; (ii) . In case (i), it follows from Lemma 4.1 that . For case (ii), it follows from Lemma 4.4 and its proof that and as locally uniformly in , which ends the proof.
5. Asymptotic profiles of spreading solutions
Throughout this section we assume that (H) holds and is a solution of () for which spreading happens. In order to determine the spreading speed, we will construct some suitable sub- and supersolutions based on semi-waves. Let and be given in Theorem 3.4. The first subsection covers the proof of the boundedness for and . Based on these results, we prove Theorem 1.4 in the second subsection.
5.1. Boundedness for and .
Let us begin this subsection with the following estimate.
Lemma 5.1**.**
Let be a solution of () for which spreading happens. Then for any , there exist small , and such that for ,
- (i)
**
- (ii)
u(t,x)\geqslant u^{*}\big{(}1-Me^{-\beta^{*}t}\big{)}\quad\mbox{for }x\in[-ct,ct];**
- (iii)
u(t,x)\leqslant u^{*}\big{(}1+Me^{-\beta^{*}t}\big{)}\quad\mbox{for }x\in[g(t),h(t)].**
Proof.
In order to prove conclusions (i) and (ii), inspired by [16], we will use the semi-wave to construct the suitable subsolution. Here we mainly use the the monotonicity and exponentially convergent of .
(i) Since is the unique positive solution of
[TABLE]
then it is easy to check that . Since for and as , thus there is such that for . Thus there exists such that and for . This means that is increasing in . Let be small. Define
[TABLE]
for and . Then is a continuous function for and , , thus there exists such that for . By continuity, there exists small such that for , . Furthermore, as , then there is a constant such that
[TABLE]
Inspired by [16], let us construct the following function:
[TABLE]
and denote and be the zero points of with , that is
[TABLE]
In the following, we will show that is a subsolution of problem (). We only prove the case where , since the other is analogous. For any function depended on , we write if no confusion arises. For simplicity of notations, we will write
[TABLE]
Firstly, a direct calculation shows that for ,
[TABLE]
Assume that , and choose large such that in . The monotonicity of and its exponential rate of convergence to at imply that if we choose sufficiently large, then there exist positive constants , and such that
[TABLE]
Set with and , where is the unique zero point of
[TABLE]
Thus, when and , since , then
[TABLE]
provided that is sufficiently large.
For the part , then for and sufficiently large , there are two positive constants and where such that , and
[TABLE]
thus we have
[TABLE]
Now let us choose satisfies
[TABLE]
with sufficiently large, and \kappa:=d_{2}e^{\beta\tau}+d\big{(}e^{\beta\tau}-1\big{)}+2\beta, then . Hence from the above we obtain that in this part.
Next, let us check the free boundary condition. When , we set and , then
[TABLE]
We differentiate (5.3) with respect to to obtain
[TABLE]
By shrinking and enlarge if necessary, then we can see that , and . This, together with (5.3), yields that . Since for and and as , thus we have
[TABLE]
Thanks to the choice of , we can compute that
[TABLE]
where is a positive constant, \kappa:=d_{2}e^{\beta\tau}+d\big{(}e^{\beta\tau}-1\big{)}+2\beta>2\beta and we have used that by shrinking if necessary, then .
It follows from (5.4), (5.5), (5.6) and the monotonicity of in that
[TABLE]
Using (5.3) again, it is easy to see that is decreasing in , thus for all ,
[TABLE]
Since is a spreading solution of (), then there exists such that
[TABLE]
Consequently, is a subsolution of problem (), then we can apply the comparison principle to conclude that , for , . This, together with (5.7), implies that
[TABLE]
with . Similarly, by enlarging if necessary, we can have for . Thus result (i) holds for large .
(ii) From the proof of (i), it is easy to see that for . The monotonicity of and its exponential rate of convergence to at can be used again to conclude that for any there exist constants , such that for any and ,
[TABLE]
Based on above results, we can find large such that for and ,
[TABLE]
where is sufficiently large and \beta^{*}:=\frac{1}{2}\min\big{\{}\nu(c^{*}-c),\ \beta,\ d-f^{\prime}(u^{*})\big{\}}. The case where can be proved by a similar argument as above. The proof of (ii) is now complete.
(iii) Thanks to the choice of the initial data, we know that for any given and ,
[TABLE]
This completes the proof. ∎
Next we prove the boundedness of and show that in the domain , where is a large number.
Proposition 5.2**.**
Assume that spreading happens for the solution . Then
- (i)
there exists such that
[TABLE]
- (ii)
for any small , there exists and such that
[TABLE]
Proof.
In order to prove conclusions in this proposition, inspired by [12], we will use the semi-wave to construct the suitable sub- and supersolution. Compared with [12], our problem deal with the case where . Due to , there will be some space-translation of the semi-wave , which make our problem difficult to deal with. To overcome this difficulty, we mainly use the the monotonicity and exponentially convergent of . Moreover, this idea also be used in Lemma 5.6. For clarity we divide the proof into several steps.
. To give some upper bounds for and .
Fix . It follows from Lemma 5.1 that there exist , , and such that for , (i), (ii) and (iii) in Lemma 5.1 hold. Thanks to (H), by shrinking if necessary, we can find small such that
[TABLE]
For any large satisfying , there is such that . Since as , we can find such that
[TABLE]
Now we construct a supersolution to () as follows:
[TABLE]
where is a positive constant to be determined below.
Clearly, for all , , \bar{u}\big{(}t,\bar{h}(t)\big{)}=0, and
[TABLE]
if we choose with . By the definition of we have for . It then follows from (5.11) that for ,
[TABLE]
which yields that for .
We now show that
[TABLE]
Thanks to the definition of and the monotonicity of in , we can find a decreasing function for , such that
[TABLE]
which implies that
[TABLE]
As , thus in what follows, we only consider the case x\in\big{[}\eta(t),\bar{h}(t)\big{]}. Set q_{\tau}:=q_{c^{*}}\big{(}\bar{h}_{\tau}-x\big{)} for convenience. A direct calculation shows that, for ,
[TABLE]
for some . Since
[TABLE]
where , there are and such that
[TABLE]
Moreover, we can compute that
[TABLE]
For any given , by enlarging if necessary, we have that
[TABLE]
When and , it then follows that
[TABLE]
provided that is sufficiently large, and we have used for , for , (5.10), (5.14) and (5.15). Thus in this case.
When and , for sufficiently large , we have
[TABLE]
where , , and (5.15) are used.
Summarizing the above results we see that is a supersolution of (). Thus we can apply the comparison principle to deduce
[TABLE]
By the definition of we see that, for , we have
[TABLE]
For any , if we choose large such that , then we have
[TABLE]
which ends the proof of Step 1.
. To give some lower bounds for and .
Let , , and be as before. By shrinking if necessary, we can find large such that
[TABLE]
We will define the following functions
[TABLE]
where is a positive constant to be determined later.
We will prove that is a subsolution to () for . Firstly, for ,
[TABLE]
Next, we check that and satisfy the required conditions at . It is obvious that . If we choose with , then
[TABLE]
Later, let us check the initial conditions. From Lemma 5.1, it is easy to see that
[TABLE]
for and .
Finally we will prove that for . Put and . It is easy to check that
[TABLE]
for some . It follows from (5.13) that there are two constants , such that
[TABLE]
Moreover, we can compute that
[TABLE]
For any given , by enlarging if necessary, we have that
[TABLE]
When and , it then follows that
[TABLE]
provided that is sufficiently large, and we have used \big{(}1-\theta_{1}Me^{-\beta^{*}(t-\tau)}\big{)}q_{\tau}\in[u^{*}-\rho,u^{*}] and (5.18) for , and (5.10), (5.19), (5.20). Thus in this case.
When and , for sufficiently large , we have
[TABLE]
where , and (5.20) are used.
Consequently, is a subsolution to (), then the comparison principle implies that
[TABLE]
which yields that
[TABLE]
where . Combining with (5.16) we obtain (5.8).
On the other hand, for any , since , there exists such that
[TABLE]
It follows from (5.21) and (5.16) that
[TABLE]
which yields that for ,
[TABLE]
Moreover, if we choose such that , then
[TABLE]
which completes the proof of Step 2.
. Completion of the proof of (5.9). Denote and , then by (5.17) and (5.22) we have
[TABLE]
This yields the estimate in (5.9), which completes the proof of this proposition. ∎
Using a similar argument as above we can obtain the following result.
Proposition 5.3**.**
Assume that spreading happens for the solution . Then
- (i)
there exists such that
[TABLE]
- (ii)
for any small , there exists and such that
[TABLE]
5.2. Asymptotic profiles of the spreading solutions
This subsection is devoted to the proof of Theorem 1.4. We will prove this theorem by a series of results. Firstly, it follows from Proposition 5.2 that there exist positive constant such that
[TABLE]
Let us use the moving coordinate and set
[TABLE]
Then solves
[TABLE]
Let be an arbitrary sequence satisfying for . Define
[TABLE]
Lemma 5.4**.**
Subject to a subsequence,
[TABLE]
where , , , and satisfies
[TABLE]
Proof.
It follows from the proof of Lemma 2.2 that there is such that for all . One can deduce that
[TABLE]
Define
[TABLE]
and direct computations yield that
[TABLE]
for , , and
[TABLE]
Since , then f\Big{(}w_{n}\Big{(}t-\tau,\frac{H_{n}(t)z+c^{*}\tau}{H_{n}(t-\tau)}\Big{)}\Big{)} is bounded. For any given and , using the partial interior-boundary estimates and the Sobolev embedding theorem (see [13, 18]), for any , we obtain
[TABLE]
where is a positive constant depending on and but independent of and . Thanks to this, we have
[TABLE]
with is a positive constant independent of and . Hence by passing to a subsequence we may assume that as ,
[TABLE]
where . Based on above results, we can see that satisfies that
[TABLE]
Define V(t,y)=W\big{(}t,\frac{y}{H(t)}\big{)}. It is easy to check that satisfies (5.27) and (5.26) holds. ∎
Later, we show by a sequence of lemmas that is a constant and hence
[TABLE]
Since for all , then for . Denote
[TABLE]
it follows from the proof of Proposition 5.2 that for and large,
[TABLE]
Letting we have
[TABLE]
Define
[TABLE]
and
[TABLE]
Then
[TABLE]
and
[TABLE]
By a similar argument as in [13], we have the following result.
Lemma 5.5**.**
, , and there exist two sequences , such that
[TABLE]
uniformly for in compact subsets of , and
[TABLE]
uniformly for in compact subsets of .
Based on Lemma 5.5, we have the following lemma.
Lemma 5.6**.**
, and hence is a constant, which yields .
Proof.
Argue indirectly we may assume that . Choose . We will show next that there is such that
[TABLE]
which implies that . This contraction would complete the proof.
To complete the proof, we need to prove that for given , there exist and such that
[TABLE]
It follows from that there exist and such that
[TABLE]
By Lemma 5.5, for any , there exist , such that for and ,
[TABLE]
Set and , then satisfies
[TABLE]
It follows from Lemma 5.5 and (5.29) that there is such that for ,
[TABLE]
Thanks to (H), for small with is given in the proof of Proposition 5.2, there is small such that
[TABLE]
and we can find independent of satisfies
[TABLE]
Let us construct the following supersolution of problem (5.30):
[TABLE]
Since \lim_{y\to-\infty}\big{(}1+N\varepsilon e^{-\beta_{0}(t-\tilde{s}_{n})}\big{)}\phi\big{(}y-\bar{G}(t)\big{)}>u^{*}, then there is a smooth function of such that as and \big{(}1+N\varepsilon e^{-\beta_{0}(t-\tilde{s}_{n})}\big{)}\phi\big{(}\bar{K}(t)-\bar{G}(t)\big{)}=u^{*}. We will check that is a supersolution for and . We note that
[TABLE]
Firstly, it follows from (5.31) that for ,
[TABLE]
In view of (5.32), we have
[TABLE]
for and . By definition and direct computation yields
[TABLE]
if we choose with . Since , it then follows from the definition of that .
Finally, let us show
[TABLE]
Put , and \phi_{\tau}:=\phi\big{(}y-\bar{G}(t-\tau)\big{)}. It is easy to compute that
[TABLE]
Since
[TABLE]
where , there are two constants and such that
[TABLE]
Moreover, we can compute that
[TABLE]
For any given , by shrinking if necessary, we have that
[TABLE]
For and , direct calculation implies
[TABLE]
provided that is sufficiently large, and we have used \big{(}1+\theta_{2}\zeta e^{\beta_{0}\tau}\big{)}\phi_{\tau}\in[u^{*}-\eta,u^{*}+\eta] for , (5.33), for , (5.35) and (5.36).
When and , for sufficiently large , we have
[TABLE]
where , , and (5.36) are used.
Thus (5.34) holds, then we can apply the comparison principle to conclude that
[TABLE]
This, together with the definition of , yields that for . By shrinking if necessary, we obtain
[TABLE]
In the following, we show for all large . As in the construction of supersolution, for any , there exists such that, for ,
[TABLE]
We also can find independent of such that
[TABLE]
We can define a subsolution as follows:
[TABLE]
Since , there are and such that for all , which implies that . Let us fix such that . By enlarging if necessary we may assume that . Denote .
By a similar argument as above and in Step 2 of Proposition 5.2, we can show that is a subsolution of problem (5.30) by taking sufficiently large. The comparison principle can be used to conclude that
[TABLE]
which implies that for . By shrinking if necessary, we have
[TABLE]
This completes the proof of this lemma. ∎
Theorem 5.7**.**
Assume that (H) and spreading happens. Then there exists such that
[TABLE]
[TABLE]
where be the unique solution of (1.13).
Proof.
It follows from Lemmas 5.4 and 5.6 that for any , by passing to a subsequence, in . The arbitrariness of implies that and as , which proves (5.40).
In what follows, we use the moving coordinate to prove (5.41). Set
[TABLE]
[TABLE]
then the pair solves
[TABLE]
By the same reasoning as in the proof of Lemma 5.4, the parabolic regularity to (5.42) plus the Sobolev embedding theorem can be used to conclude that, by passing to a further subsequence if necessary, as , in , and satisfies, in view of ,
[TABLE]
This is equivalent to (5.27) with and . Hence we can conclude
[TABLE]
Thus we have proved that, as ,
[TABLE]
This, together with the arbitrariness of , yields that
[TABLE]
Then, for any ,
[TABLE]
Using the limit as we obtain
[TABLE]
Finally we prove (5.41). For any given small , it follows from (5.9) in Proposition 5.2 that there exist two positive constants and such that
[TABLE]
Since as , there exists such that
[TABLE]
Taking large such that for , then we obtain
[TABLE]
Taking in (5.43) we see that for some , we have
[TABLE]
This completes the proof of (5.41). ∎
Taking use of a similar argument as above one can obtain the following result.
Theorem 5.8**.**
Assume that (H) and spreading happens. Then there exists such that
[TABLE]
[TABLE]
where be the unique solution of (1.13).
Proof of Theorem 1.4. The results in Theorem 1.4 follow from Theorems 5.7 and 5.8.
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