The peloton superorganism and protocooperative behavior
Hugh Trenchard

TL;DR
This paper introduces a theoretical framework for protocooperative behavior in pelotons, modeling the transition between cooperation and free-riding based on drafting benefits, cyclists' power, and speed, highlighting superorganism-like properties.
Contribution
It presents a novel model of protocooperative behavior in pelotons, defining thresholds and characteristics that distinguish cooperation from free-riding, with implications for biological systems.
Findings
Identifies a drafting coefficient threshold for cooperative behavior
Describes phases of low-speed sharing and free-riding in pelotons
Suggests similar behaviors in other energy-saving biological systems
Abstract
A theoretical framework for protocooperative behavior in pelotons (groups of cyclists) is proposed. A threshold between cooperative and free-riding behaviors in pelotons is modeled, together comprising protocooperative behavior (different from protocooperation). Protocooperative behavior is a function of: 1. two or more cyclists coupled by drafting benefit, 2. cyclists' current power output or speed, and 3. cyclists' maximal sustainable outputs (MSO). Characteristics of protocooperative behavior include: 1. relatively low speed phase in which cyclists naturally pass each other and share highest-cost front positions, and 2. free-riding phase in which cyclists maintain speeds of those ahead, but cannot pass. Threshold for protocooperative behavior is equivalent to coefficient of drafting d, below which cooperative behavior occurs; above which free-riding occurs up to a second threshold…
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